Do you drink coffee? A lot of people do—coffee is a major tropical crop, valued at over \$100 billion per year. The most popular species of coffee, Coffea arabica, originated in Africa and is now planted extensively in Central and South America. The pest called the coffee berry borer beetle (Hypothenemus hampeii) moved along with the coffee and is a major problem for coffee farmers, sometimes destroying half of the coffee crop in mature plantations.
Look at the graph. Why did the researchers bother to collect data from the coffee shrubs before adding the nets to treatment plots to exclude birds?

Do you drink coffee? A lot of people do—coffee is a major tropical crop, valued at over \$100 billion per year. The most popular species of coffee, Coffea arabica, originated in Africa and is now planted extensively in Central and South America. The pest called the coffee berry borer beetle (Hypothenemus hampeii) moved along with the coffee and is a major problem for coffee farmers, sometimes destroying half of the coffee crop in mature plantations. Draw a simple model to compare the relative species richness and species diversity of these two coffee plantations: one is clear-cut and planted with only coffee shrubs, the other retains some natural forest. For symbols, use a few letters to represent different species of plants and a few numbers to represent different species of animals in a square 1-ha plot.

Do you drink coffee? A lot of people do—coffee is a major tropical crop, valued at over \$100 billion per year. The most popular species of coffee, Coffea arabica, originated in Africa and is now planted extensively in Central and South America. The pest called the coffee berry borer beetle (Hypothenemus hampeii) moved along with the coffee and is a major problem for coffee farmers, sometimes destroying half of the coffee crop in mature plantations.

Based on what you have learned in this chapter about global trends in terrestrial ecosystems, what is currently the largest threat to the wet tropical forests in Central America?

a. Habitat destruction, such as the clearing of forests for coffee plantations

b. Exotic species, such as the introduction of coffee berry borers

c. Pollution, such as the use of pesticides in coffee plantations

d. Climate change, such as changes in precipitation patterns

Do you drink coffee? A lot of people do—coffee is a major tropical crop, valued at over \$100 billion per year. The most popular species of coffee, Coffea arabica, originated in Africa and is now planted extensively in Central and South America. The pest called the coffee berry borer beetle (Hypothenemus hampeii) moved along with the coffee and is a major problem for coffee farmers, sometimes destroying half of the coffee crop in mature plantations.

Pesticides are generally ineffective in killing coffee berry borers. Ecologist Daniel Karp and colleagues conducted a study in Costa Rica to determine if natural predators like birds reduce the beetle population. The researchers selected 12 control plots and used nets to exclude birds from 12 similar treatment plots. What is the take-home message of the graph? (Remember, * means P < 6 0.05)

Do you drink coffee? A lot of people do—coffee is a major tropical crop, valued at over \$100 billion per year. The most popular species of coffee, Coffea arabica, originated in Africa and is now planted extensively in Central and South America. The pest called the coffee berry borer beetle (Hypothenemus hampeii) moved along with the coffee and is a major problem for coffee farmers, sometimes destroying half of the coffee crop in mature plantations. The abundance of birds and other predators of borer beetles depends on how much natural forest is left within and around the coffee plantations. What is the approximate percentage increase in borer predators per hectare if forest cover is doubled from 15 to 30 percent?