Which sequence correctly outlines the steps of the malate-aspartate shuttle?

Oxaloacetate is converted to aspartate, aspartate enters the matrix, aspartate is oxidized to malate, malate is reduced to oxaloacetate.

Malate is oxidized to oxaloacetate, oxaloacetate enters the matrix, oxaloacetate is reduced to malate, malate is converted to aspartate.

Aspartate is converted to oxaloacetate, oxaloacetate enters the matrix, oxaloacetate is reduced to malate, malate is oxidized to aspartate.

Oxaloacetate is reduced to malate, malate enters the matrix, malate is oxidized to oxaloacetate, oxaloacetate is converted to aspartate.