뒤로Life of a Flowering Plant: Structure, Growth, and Reproduction
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Flowering Plant Structure
Main Systems of a Flowering Plant
Flowering plants are composed of two main systems: the root system (below ground) and the shoot system (above ground). Each system is specialized for distinct functions essential to plant survival and reproduction.
Shoot system: Collects light, performs photosynthesis, enables reproduction, and transports nutrients between leaves and roots.
Root system: Acquires water and nutrients, anchors the plant, and stores starch.

Key Structures in the Shoot System
Apical meristems: Regions at the tips of roots and shoots where growth occurs.
Petiole: Stalk attaching the leaf blade to the stem.
Blade: The broad, flat part of the leaf (not labeled in image).
Flower: Non-photosynthetic structure containing reproductive organs.
Internode: Space between adjacent leaves.
Bud: Growth area for stems, leaves, and flowers; terminal (top) or lateral (side).
Stem: Supports the plant upright.
Plant Growth and Meristems
Types of Growth
Plant growth occurs at specialized regions called meristems, which contain undifferentiated cells capable of division. These cells divide by mitosis, and as they mature, they differentiate to perform specific functions.
Primary growth: Increases plant height and develops specialized structures; occurs at apical meristems.
Secondary growth: Increases girth of stems and roots; occurs at lateral meristems.
Herbaceous plants: Only primary growth; flexible stems; annuals (live one year).
Woody plants: Both primary and secondary growth; perennials (live many years).
Plant hormones regulate growth, fruit ripening, and flower scent production, and are transported in the phloem.
Secondary Growth and Annual Rings
Woody plants exhibit secondary growth, producing annual rings that record each year of growth. The structure of a woody stem includes:
Heartwood: Older xylem, no longer conducts water, stores metabolic wastes.
Sapwood: Younger xylem, still conducts water.
Bark: Tough outer layer protecting inner tissues.
Light-colored rings form in spring/early summer (abundant water), dark rings in late summer/fall (less water). One light and one dark ring represent one year.

Plant Tissue Types
Three Major Tissue Types
Dermal tissue: Covers the plant's surface; includes the epidermis (outer layer) and peridermis (replaces dead epidermal cells). The epidermis is covered by a waxy cuticle to prevent water loss.
Ground tissue: Makes up most of the plant body; involved in photosynthesis, support, and storage.
Vascular tissue: Pipeline system for transporting water, minerals, and nutrients throughout the plant.
Ground Tissue Cell Types
Parenchyma cells: Thin-walled, alive, involved in photosynthesis, hormone secretion, support, and food storage.
Collenchyma cells: Elongated, alive, provide support for young and herbaceous plants.
Sclerenchyma cells: Thick-walled, dead, provide support for adult plants and fruit structures.
Vascular Tissue: Xylem and Phloem
Xylem: Conducts water and minerals upward from roots; composed of sclerenchyma fibers, tracheids, and vessel elements.
Phloem: Conducts water, sugars, amino acids, and hormones throughout the plant; composed of sieve-tube elements and companion cells.
Xylem Structure
Sclerenchyma fibers: Dead cells with lignin for support.
Tracheids: Needle-like, dead cells forming tubes for water movement.
Vessel elements: Wide, dead cells forming continuous tubes for water transport.

Phloem Structure
Sieve-tube elements: Pass sugars and organic molecules; separated by sieve plates; alive but not metabolically active.
Companion cells: Regulate and nourish sieve-tube elements.

Leaf Structure and Function
Leaf Anatomy
Leaves are specialized for photosynthesis and gas exchange. Key structures include:
Cuticle: Waxy, waterproof layer preventing water loss.
Stoma (stomata): Pores for gas exchange; CO2 in, O2 and H2O vapor out.
Guard cells: Open and close stomata by changing turgor pressure.
Bundle-sheath cell: Surrounds vascular bundles (veins).
Mesophyll: Interior leaf tissue; palisade layer (photosynthesis) and spongy layer (gas exchange).

Stem and Root Anatomy
Stem Structure
Pith: Central part; supports, stores food, may photosynthesize; made of parenchyma cells.
Cortex: Between pith and epidermis; similar function to pith.
Epidermis: Outer protective layer.
Vascular bundle: Contains xylem and phloem.
Vascular cambium: Meristematic tissue forming new xylem and phloem.
Root System and Anatomy
Taproot: Main root for anchorage and storage.
Branch roots and root hairs: Increase surface area for absorption.
Root cap: Protects growing root tip and secretes lubricant.
Fibrous roots: Found in grasses and palms; all roots similar in size.

Root Anatomy
Epidermis: Outer protective layer.
Cortex: Stores starch; loosely packed cells.
Endodermis: Closely packed cells surrounding the vascular cylinder.
Vascular cylinder: Contains xylem and phloem; pericycle is the layer closest to endodermis, conducting water and nutrients inward.
Casparian strip: Waxy layer in endodermal cells, forcing water into the vascular cylinder.

Plant Root Symbioses
Mutualistic Relationships
Fungal mycorrhizae: Fungi that help plants acquire nutrients by releasing enzymes to weather rocks and make minerals available.
Nitrogen-fixing bacteria: Bacteria in root nodules convert atmospheric nitrogen into ammonium or nitrate ions usable by plants (e.g., legumes such as peas and beans).

Monocots vs. Dicots (Eudicots)
Key Differences
Feature | Monocots | Dicots (Eudicots) |
|---|---|---|
Vascular bundles | Scattered | In a ring |
Root type | Fibrous | Taproot |
Cotyledons | 1 | 2 |
Leaf veins | Parallel | Netted |
Flower parts | Multiples of 3 | Multiples of 4 or 5 |
In angiosperms, the sporophyte is the dominant form, while the gametophyte is small and grows within the sporophyte.
Flower Anatomy and Reproduction
Flower Structure
Stamen: Male reproductive structure (anther and filament).
Anther: Produces pollen.
Filament: Supports the anther.
Carpel: Female reproductive structure (stigma, style, ovary).
Ovary: Produces ovules.
Stigma: Receives pollen.
Sepal: Supports the flower bud.
Petal: Pigmented, attracts pollinators.

Complete vs. Incomplete Flowers
Complete flowers: Contain all four parts (sepals, petals, stamens, carpels).
Incomplete flowers: Lack one or more parts; can be perfect (both stamens and carpels) or imperfect (only stamens or only carpels).

Male Gametophyte: Pollen Formation
Microspore mother cell (MMC) develops in pollen sacs.
MMC undergoes meiosis to form four haploid microspores.
Each microspore divides mitotically to form an immature pollen grain.
The generative cell divides to form two sperm cells, creating a mature pollen grain.

Female Gametophyte: Embryo Sac Formation
Megaspore mother cell (MMC) develops in the ovule.
MMC undergoes meiosis to form four haploid megaspores; three degenerate.
The remaining megaspore divides mitotically three times to form eight nuclei in one cell.
Cytokinesis forms seven cells (embryo sac); one is the egg cell.

Pollination and Double Fertilization
Pollination occurs when a pollen grain lands on the stigma and grows a pollen tube to the ovary. Double fertilization involves:
One sperm fertilizes the egg, forming the zygote.
The other sperm fuses with two polar nuclei, forming the triploid endosperm (nutrient tissue for the embryo).

Fruit and Seed Development
After fertilization, the ovary develops into a fruit, which protects the seeds and aids in their dispersal. Fruits can be fleshy (eaten by animals) or dry (dispersed by wind or attachment to animals).

Seed Dispersal Mechanisms
Wind: Seeds adapted for wind dispersal are lightweight or have structures for floating in the air.
Water: Some seeds float and are dispersed by water.
Animals: Seeds may be eaten and excreted, or attach to animal fur for transport.

Seed Germination
Requirements and Process
Requires moisture and appropriate temperature.
Some seeds need to be dried, exposed to cold, or have their seed coat broken to germinate.
Many seeds undergo dormancy (reduced metabolic activity) before germination.
In monocots, the shoot is protected by a sheath called the coleoptile.
In dicots, the shoot lacks a coleoptile; the region above the cotyledons is the epicotyl, below is the hypocotyl.
In dicots, the endosperm is used up by the growing cotyledons.