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Introduction to Ecology and Behavioural Ecology: Study Guide

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Introduction to Ecology

What is Ecology?

Ecology is the scientific study of how organisms interact with each other and with their environment. The primary goals of ecology are to understand the distribution and abundance of organisms and to identify the impacts humans have on the environment. Ecology is closely linked with evolution, as evolutionary processes shape ecological interactions and vice versa.

  • Distribution: Where organisms are found geographically.

  • Abundance: How many individuals of a species are present in a given area.

  • Human Impact: Human activities can alter ecological systems, sometimes with unintended consequences.

Cover of Silent Spring by Rachel Carson

Example: The publication of Silent Spring by Rachel Carson in 1962 highlighted the ecological consequences of DDT pesticide use, which led to declines in bird populations due to bioaccumulation and biomagnification.

DDT and bioaccumulation in a food chain

Levels of Ecological Study

Ecological Scales

Ecology can be studied at multiple biological scales, each focusing on different aspects of the interactions between organisms and their environment:

  • Biosphere: Global processes and patterns, such as climate and ocean currents, that affect life on Earth.

  • Ecosystem: The flow of energy and matter between living (biotic) and non-living (abiotic) components.

  • Community: Interactions among different species living in the same area.

  • Population: Dynamics of a single species, including changes in size, density, and structure over time and space.

  • Individual: How the morphology, physiology, and behavior of an organism allow it to survive and reproduce in its environment.

Diagram of ecological scales from individual to biosphere

Determinants of Distribution and Abundance

Biogeography and Limiting Factors

Biogeography is the study of how organisms are distributed geographically. The distribution and abundance of species are influenced by both abiotic (non-living) and biotic (living) factors.

  • Abiotic Factors: Climate (temperature and moisture), soil, light, and nutrients.

  • Biotic Factors: Interactions with other species, such as competition, predation, mutualism, and parasitism.

Global distribution of animals and plants

Niche Models: These models show the range of environmental conditions a species can tolerate. Some species have broad tolerances (generalists), while others are more specialized (specialists).

Niche models showing range of conditions for two microbes

Climate and Its Effects on Ecology

Latitudinal and Seasonal Variation

Climate, defined by long-term patterns of temperature and precipitation, is a major determinant of species distribution. Solar radiation per unit area declines with increasing latitude, making the tropics warmer and the poles colder.

Solar radiation per unit area declines with latitude

Global air circulation patterns, such as Hadley cells, create moist tropics and dry subtropical regions.

Hadley cells and global air circulation

Seasonality is caused by the Earth's 23.5° tilt on its axis, resulting in varying solar input throughout the year.

Earth's orbit and tilt create seasons

Regional effects, such as rain shadows created by mountains and oceanic moderation of climate, further influence local climates.

Rain shadow effect of mountains Ocean gyres and climate moderation

Terrestrial Biomes

Major Biome Types

Terrestrial biomes are major groupings of plant and animal communities defined by dominant vegetation and governed by average temperature and precipitation.

Map of major terrestrial biomes

Biome

Climate

Key Features

Tropical Wet Forest

Warm, high moisture

High diversity, high productivity

Subtropical Desert

Extreme temperatures, low moisture

Low diversity, adaptations for drought

Temperate Grassland

Moderate temp, low-moderate moisture

Fertile soil, fire regimes, dense growth

Temperate Forest

Moderate temp, moderate moisture

Deciduous trees, seasonal photosynthesis

Temperate Coniferous Forest

Mild winters, high moisture

Conifer canopy, diverse understory

Boreal Forest (Taiga)

Low temp, low moisture

Cold-tolerant conifers, low diversity

Arctic Tundra

Very low temp, very low moisture

Low productivity, permafrost

Tropical wet forest Subtropical desert

Aquatic Biomes

Physical Factors Affecting Aquatic Life

About 70% of Earth's surface is covered by water. The distribution and abundance of aquatic organisms are influenced by four main physical factors:

  • Salinity: The concentration of dissolved solutes; affects osmoregulation in organisms.

  • Water Flow: Movement of water presents physical challenges and affects dispersal.

  • Nutrient Availability: Essential for productivity; affected by processes like lake turnover.

  • Water Depth: Dictates light penetration and thus photosynthetic activity.

Example: Lake turnover redistributes nutrients and oxygen, supporting aquatic life.

Introduction to Behavioural Ecology

Behaviour and Adaptation

Behavioural ecology studies how behavioural adaptations evolve in response to ecological selection pressures, affecting an organism's fitness. Behaviour is defined as an action in response to a stimulus and can be analyzed at two levels:

  • Proximate (Mechanistic) Causation: Explains how actions occur (e.g., genetic, neurological, hormonal mechanisms).

  • Ultimate (Evolutionary) Causation: Explains why actions occur (e.g., evolutionary history, fitness benefits).

Example: Male starlings sing in spring due to hormonal changes (proximate) and to attract mates (ultimate).

Common Questions in Behavioural Ecology

  1. What should I eat?

  2. Where should I live?

  3. Whom should I mate with?

  4. How should I communicate?

  5. When should I cooperate?

Foraging Behaviour

Foraging behaviour is influenced by genetic variation and fitness trade-offs. For example, in Drosophila larvae, "rovers" and "sitters" exhibit different foraging strategies based on population density, controlled by the for gene.

  • Optimal Foraging Theory: Predicts that animals maximize benefits and minimize costs when foraging.

Habitat Selection and Navigation

Animals select habitats based on quality, population density, and competition. Navigation strategies include:

  • Piloting: Using landmarks.

  • Compass Orientation: Using the sun, stars, or Earth's magnetic field.

  • True Navigation: Locating a specific place on Earth.

Migration is a costly behaviour but may increase survival and reproductive success by tracking seasonal resources.

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