Campbell's Biology, 13th edition

Published by Pearson (April 9, 2026) © 2027
  • Lisa A. Urry
  • Peter V. Minorsky
  • Kerry L Hull
  • Rebecca B. Orr

Title overview

For courses in general biology.

Excellence. Accuracy. Innovation.

Campbell Biology delivers a proven, current, absorbing experience that guides students to a true understanding of biology. The author team advances Neil Campbell's vision: equip students at their individual skill levels with tools, visuals, resources, and activities to engage them in learning.

The 13th Edition offers new Practice Applying Your Learning question sets in each unit. A wealth of new and updated content includes updates in gene editing and stem cell research; artificial intelligence and its role in bioinformatics and medical diagnosis; global climate and population changes; and much more. The trusted approach and latest enhancements of this leading text are also well aligned with the Vision and Change core competencies.

Hallmark features of this title

  • Make Connections figures link content from different chapters to represent big-picture relationships visually.
  • Make Connections questions show the connections between areas of biology by relating chapter content to material presented earlier.
  • Summary of Key Concepts questions at the end of each chapter reinforce the main ideas in each chapter.
  • Test Your Understanding questions prepare students for exam-type questions and encourage higher-level thinking.
  • Scientific Skills Exercises use real data to help build skills in data analysis, graphing, experimental design, and math.
  • Problem-Solving Exercises guide students to apply scientific skills and interpret real data in the context of a real-world problem.

New and updated features of this title

  • NEW/UPDATED content throughout includes:
    • updates in gene editing, stem cell research, genomics, cancer genetics, biologic drugs, and biomaterials (Chapters 17, 18, 20)
    • introduction of artificial intelligence and its role in determining protein structure, bioinformatics, and medical diagnosis (Chapters 1, 5, 11, 21)
    • updated coverage of the evolutionary origins of eukaryotes and the phylogeny of archaea, unicellular eukaryotes, plants, and animals (Chapters 26 - 29, 32, 33)
    • SARS-CoV-2 and the COVID-19 pandemic, and expanded discussions of emerging and zoonotic pathogens (Chapters 19, 43, 54)
    • updated data on global climate and population changes (Chapters 3, 55, 56).
  • NEW Practice Applying Your Learning question sets in each unit: Each set offers a short biological scenario followed by multiple True/False questions at lower and higher Bloom’s levels. An alternate version of each Practice Applying Your Learning question set is available in Mastering Biology; instructors can assign 1 set for practice or formative assessment, and an alternate version for summative assessment.

Key features

Features of Mastering Biology for the 13th Edition

  • NEW: An alternate version of each Practice Applying Your Learning question set from the textbook is available in Mastering. This enables instructors to assign 1 set for practice or formative assessment, and then an alternate version for summative assessment.
  • NEW: Interactive reading assignments let students read, watch, and practice in one seamless experience. Instructors can now assign specific sections of the eTextbook plus coaching and drag and drop assignments, Dynamic Study Modules, and more for auto-grading.
  • Dynamic Study Modules pose a handful of questions and then respond to each student’s progress in real time. Learners deepen their grasp of concepts as they go.
  • Interactive Labs: A comprehensive virtual lab experience engages students in learning to think like scientists. Built around the process of science, these online labs feature real-world scenarios and guided feedback.
  • Study Area: Abundant online self-study resources are available to students, including Active Reading Guides, Figure Walkthroughs, videos, animations, Get Ready for This Chapter, Practice Tests, Cumulative Test, and more.
  • Instructor Resources Area: 1-stop shopping for PowerPoints, Clicker Questions, animations, videos, the Test Bank, and more.

Features of Pearson+ eTextbook for the 13th Edition

  • NEW: AI-Powered Study Tool pulls from vetted Campbell Biology content to help students achieve proficiency and master key course concepts. Support is provided anytime, anywhere within the assigned textbook through instant explanations, on-demand summaries, and practice questions with feedback.
  • EXPANDED: Interactive CheckPoint Questions are embedded at point-of-use. These short sequences of interactive questions diagnose and remediate misconceptions around topics that are common “stumbling blocks” to understanding.
    • Customized feedback guides students through an evidence-based approach to reason their way to learning and retaining the content.
    • The CheckPoints are also assignable in Mastering Biology.
  • Popular Figure Walkthroughs guide students through key figures with narrated explanations and figure mark-ups that reinforce important points.
  • AAAS Science in the Classroom website links provide research papers from Science magazine, with annotations to help students understand content.

Table of contents

  • 1. Evolution, the Themes of Biology, and Scientific Inquiry

UNIT 1: THE CHEMISTRY OF LIFE

  • Interview: Kristi Anseth
  • 2. The Chemical Context of Life
  • 3. Water and Life
  • 4. Carbon and the Molecular Diversity of Life
  • 5. The Structure and Function of Large Biological Molecules

UNIT 2: THE CELL

  • Interview: Julie Theriot
  • 6. A Tour of the Cell
  • 7. Membrane Structure and Function
  • 8. Introduction to Metabolism
  • 9. Cellular Respiration and Fermentation
  • 10. Photosynthesis
  • 11. Cell Communication
  • 12. The Cell Cycle

UNIT 3: GENETICS

  • Interview: Pardis Sabeti
  • 13. Meiosis and Sexual Life Cycles
  • 14. Mendel and the Gene Idea
  • 15. The Chromosomal Basis of Inheritance
  • 16. he Molecular Basis of Inheritance
  • 17. Gene Expression: From Gene to Protein
  • 18. Regulation of Gene Expression
  • 19. Viruses
  • 20. DNA Tools and Biotechnology
  • 21. Genomes and Their Evolution

UNIT 4: MECHANISMS OF EVOLUTION

  • Interview: Neil H. Shubin
  • 22. Descent with Modification: A Darwinian View of Life
  • 23. The Evolution of Populations
  • 24. The Origin of Species
  • 25. The History of Life on Earth

UNIT 5: THE EVOLUTIONARY HISTORY OF BIOLOGICAL DIVERSITY

  • Interview: Casey Dunn
  • 26. Phylogeny and the Tree of Life
  • 27. Bacteria and Archaea
  • 28. The Origin and Diversification of Eukaryotes
  • 29. Plant Diversity I: How Plants Colonized Land
  • 30. Plant Diversity II: The Evolution of Seed Plants
  • 31. Fungi
  • 32. An Overview of Animal Diversity
  • 33. An Introduction to Invertebrates
  • 34. The Origin and Evolution of Vertebrates

UNIT 6: PLANT FORM AND FUNCTION

  • Interview: Ernesto Gianoli
  • 35. Vascular Plant Structure, Growth, and Development
  • 36. Resource Acquisition and Transport in Vascular Plants
  • 37. Soil and Plant Nutrition
  • 38. Angiosperm Reproduction and Biotechnology
  • 39. Plant Responses to Internal and External Signals

UNIT 7: ANIMAL FORM AND FUNCTION

  • Interview: Adrienne Bratcher
  • 40. Basic Principles of Animal Form and Function
  • 41. Animal Nutrition
  • 42. Circulation and Gas Exchange
  • 43. The Immune System
  • 44. Osmoregulation and Excretion
  • 45. Hormones and the Endocrine System
  • 46. Animal Reproduction
  • 47. Animal Development
  • 48. Neurons, Synapses, and Signaling
  • 49. Nervous Systems
  • 50. Sensory and Motor Mechanisms
  • 51. Animal Behavior

UNIT 8: ECOLOGY

  • Interview: Elisa Bonnacorso
  • 52. An Introduction to Ecology and the Biosphere
  • 53. Population Ecology
  • 54. Community Ecology
  • 55. Ecosystems and Restoration Ecology
  • 56. Conservation Biology and Global Change

APPENDICES

  • A. Answers
  • B. Classification of Life
  • C. A Comparison of the Light Microscope and the Electron Microscope
  • D. Scientific Skills Review
  • E. Word Roots, Prefixes, and Suffixes for Learning Vocabulary

Author bios

About our authors

Lisa A. Urry (Chapter 1 and Units 1, 2, and 3) is Professor of Biology at Mills College at Northeastern University in Oakland, California. After earning a BA at Tufts University, she completed her PhD at the Massachusetts Institute of Technology (MIT). Lisa has conducted research on gene expression during embryonic and larval development in sea urchins. Deeply committed to promoting opportunities in science for women and underrepresented groups, she has taught courses ranging from introductory and developmental biology to an immersive course on the U.S./Mexico border.

Peter V. Minorsky (Unit 6) is Professor of Biology at Mercy University in New York, where he teaches introductory biology, ecology, and botany. He received his AB from Vassar College and his PhD from Cornell University. Peter has also taught at Kenyon College, Union College, Western Connecticut State University, and Vassar College. His research interests concern how plants sense environmental change. Peter received the 2008 Award for Teaching Excellence at Mercy.

Kerry L. Hull (Unit 7) is the Associate Vice-Principal, Research; Dean of Natural Science and Mathematics; and a Professor in the Department of Biology and Biochemistry at Bishop’s University in Quebec, Canada. Kerry holds a BSc in Biology and a PhD in Physiology (Endocrinology), both from the University of Alberta (Canada). She teaches physiology and anatomy courses to undergraduates, using guided inquiry and case studies as her primary teaching approaches. A comparative endocrinologist by training, her current research focuses on the development and implementation of active learning strategies in anatomy and physiology courses.

Rebecca B. Orr (Assessment Lead, eTextbook Media Integration, and Active Learning Modules) is Professor of Biology at Collin College in Plano, Texas, where she teaches introductory biology. She earned her BS from Texas A&M University and her PhD from University of Texas Southwestern Medical Center at Dallas. Rebecca has a passion for investigating strategies that result in more effective learning and retention, and she is a certified Team-Based Learning Collaborative Trainer Consultant. She enjoys focusing on the creation of learning opportunities that both engage and challenge students.

Neil A. Campbell (1946–2004) earned his MA from the University of California, Los Angeles, and his PhD from the University of California, Riverside. His research focused on desert and coastal plants. Neil’s 30 years of teaching included introductory biology courses at Cornell University, Pomona College, and San Bernardino Valley College, where he received the college’s first Outstanding Professor Award. He was also a visiting scholar at the University of California, Riverside. Neil was the founding author of Campbell Biology.

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