As a result of proofreading by DNA polymerases, the overall error rate in the completed DNA molecule is approximately __________.

• A. 1 error per 100 nucleotides
• B. 1 error per 1,000 nucleotides
• C. 1 error per 1,000,000 nucleotides
• D. 1 error per 1,000,000,000 nucleotides
• E. 1 error per 10,000,000,000 nucleotides